The squares hold the plastic cases that ¼" endmills commonly come in. There are holes for “naked” ¼" and 1/8" endmills. The slot on the right is for a collet wrench (fits the OpenBuilds one for sure). The larger holes loosely hold empty ER11 collets.
I cut this out of MDF on my CNC, but 3D printing is an option as well. If your printer prints small, not-so-round holes, make sure you scale the entire thing up in X and Y by a few percent to allow for clearance.
The author hasn't provided the model origin yet.